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franzae

Annotated Checklist of
the New World
Lucanidae

(Coleoptera: Scarabaeoidea)

VERSION 4.0 uploaded
12 October, 2013


M.J. Paulsen

Erichius franzae (Weinreich)
Photo by M.J. Paulsen.


The stag beetle family Lucanidae includes four subfamilies, 109 genera and around 1500 species worldwide (Benesh 1960; Maes 1992). In the New World there are 42 genera and 232 species with representatives of each of the four subfamilies. Most New World taxa are Neotropical; in the Nearctic region there are only 26 species in eight genera.

Traditionally, the family has been characterized as moderate to large (8-60 mm), hornless beetles possessing enlarged male mandibles and elbowed, clubbed antennae. The antennae are often stated to have an elongate first segment, and club segments that cannot be closed tightly together (White 1983; Lawrence and Britton 1991; Triplehorn and Johnson 2005).

However, there are taxa that resist all attempts to neatly characterize the family. In some species, the mandibles of the male and female are of similar size and are indistinguishable. Only one subfamily (Lucaninae) possesses antennae that are elbowed. In some aesalines, the first antennal segment is not elongated, and the antennal club segments can be closed (if loosely), prompting some authors to treat them as members of the Trogidae (Scarabaeoidea e.g., Kikuta 1986). Still other aesalines have barely asymmetrical clubs. Additionally, males of the genus Sinodendron (Syndesinae) possess both a clypeal horn and a horn-like pronotal tubercle.

Exceptions aside, almost all stag beetles possess 10-segmented antennae, with the distal 3-7 segments elongated asymmetrically into a serrate, lamellate, or flabellate club that cannot be closed tightly as in many other scarabaeoids. Despite the difficulty in characterizing them, many stag beetles are immediately recognizable and are frequently collected. Their popularity with collectors has probably led to an excess of generic, specific, and subspecific names. Yet, little comprehensive taxonomic work has been conducted on the family, and revisions and keys are needed for most genera. Currently, the only available method for identifying many species is a richly illustrated picture book (Fujita 2010), although it contains numerous misidentifications of the New World fauna and should be used with care.

The higher-level classification of the family is in need of rigorous study. The number of recognized subfamilies has varied greatly (Didier and Séguy 195; Benesh 1960; Howden and Lawrence 1974). Subfamilial placement in this checklist follows the work of Holloway (1960, 1968, 1969, 1997, 1998). The classification of the family at the tribal level is ongoing and recent changes are incorporated below (Paulsen 2010). As noted by Smith (2006), numerous tribal names in the Lucanidae were not formed correctly according to ICZN Article 13.1 (first used after 1930 but not accompanied by a description or definition) and are not available names. These include: Aegini (Maes 1992b), Allotopini (Maes 1992a), Casignetini (Maes 1992b), Chalcodinae (Didier and Séguy 1953, Lacroix 1979), Colophonini (Maes 1992b), Cyclommatini (Maes 1992a), Homoderini (Maes 1992b), Neoprosopocoilini (Maes 1992b) (which is also permanently invalid because it is not based on an available generic name), Nigidiini (Benesh 1960), Prosopocoilini (Benesh 1960), Pseudodorcini (Benesh 1960), and Rhyssonotini (Benesh 1960).

For utility and completeness, known errors in use of authorship, spelling, homonymy, and synonymy are being continued in this list because the appropriate corrections have not yet been published. Select a genus in the table below, or scroll down to view the entire checklist.

 

AESALINAE
 
SYNDESINAE
 
LAMPRIMINAE

LUCANINAE
Aegognathus   Caenolethrus   Erichius  
Altitatiayus Cantharolethrus Incadorcus
Andinolucanus   Casignetus    
Apterocyclus   Charagmophorus   Lucanus  
Apterodorcus Chiasognathus
Arnaudius Chileistomus
 
Auxicerus Dorcus
   
Brasilucanus Montesinus
 

New World Lucanidae

AESALINAE MacLeay 1819: 102
AESALINI MacLeay 1819: 102
  Lucanobium Howden & Lawrence 1974: 1507 info  
    Lucanobium guianense Paulsen 2011: 2 info French Guiana
    Lucanobium squamosum Howden & Lawrence 1974: 1508 info Venezuela
       
  Trogellus Paulsen 2013: 6  
    Trogellus catrachitus Paulsen 2013: 16 Honduras
    Trogellus chapinitus Paulsen 2013: 11 Guatemala, Honduras
    Trogellus hawksi Paulsen 2013: 12 Guatemala, Mexico
    Trogellus herrmanni Paulsen 2013: 7 Costa Rica, Panama
    Trogellus maesi Paulsen 2013: 13 Nicaragua
    Trogellus narizotus Paulsen 2013: 16 Guatemala
    Trogellus neotropicalis (Bates 1886: 2) Guatemala
    Trogellus ticiticus Paulsen 2013: 16 Costa Rica
    Trogellus trifinius Paulsen 2013: 17 El Salvador
    Trogellus trogoides (Albers 1883: 228) Mexico
      syn. Aesalus smithi Bates 1889: 382  
       
       
CERATOGNATHINI Sharp 1899: 194
  Hilophyllus Paulsen & Mondaca 2006: 6  
    Hilophyllus argentinensis (Martínez 1981: 13) (Ceratognathus) Argentina, Chile
    Hilophyllus martinezi Paulsen & Mondaca 2006: 16 Chile
    Hilophyllus penai (Martínez 1976: 101) (Ceratognathus) Chile
       
NICAGINI LeConte 1861: 130
  Nicagus LeConte 1861:130 info  
    Nicagus obscurus (LeConte 1848: 86) (Ochodaeus) Eastern USA, Canada
    Nicagus occultus Paulsen & Smith 2005: 48  Western USA: TX
         
SYNDESINAE MacLeay 1819: 103
CERUCHINI Jacquelin du Val & Fairmaire 1859: 4
  Ceruchus MacLeay 1819: 115  
    syn. Tarandus Dejean 1833: 174  
    Ceruchus piceus (Weber 1801: 84) (Lucanus) info Eastern North America
      syn. Platycerus balbi Laporte 1840: 174  
      syn. C. virginiensis Casey 1914: 372  
      nom. nud. Tarandus americanus Dejean 1833: 174  
      nom. nud. C. frontosus Knoch in Burmeister 1847: 329  
      nom. nud. C.quercicola Melsheimer in Sturm 1843: 136  
    Ceruchus punctatus LeConte 1869: 377 info Western USA, Canada
    Ceruchus striatus LeConte 1859: 85 info Western USA, Canada
       
SINODENDRINI Mulsant 1842: 600
  Sinodendron Hellwig 1792: 391  
    Sinodendron rugosum Mannerheim 1843: 262 Western USA, Canada
       
SYNDESINI MacLeay 1819: 103
  Psilodon Perty 1830: 53  
    syn. Hexaphyllum Gray 1832: 536  
    Psilodon aequinoctiale (Buquet 1840: 375) (Hexaphyllum) Colombia
       syn. H. westwoodi Hope 1840: 11  
    Psilodon buhrnheimi Grossi & Aguiar 2014: 84 Brazil (Amazonas)
    Psilodon gilberti Boucher 1993: 422 Bolivia
    Psilodon luki (Onore, Bartolozzi, and Zilioli 2011: 43) Ecuador (Pichincha)
    Psilodon schuberti Perty 1830: 54 Argentina, Brazil
      syn. H. brasiliense Gray 1832: 536  
    Psilodon paschoali Pardo-Locarno & Ríos-Málaver 2012: 247 Colombia
    Psilodon seguyi (Didier 1929: 109) (Hexaphyllum) Ecuador (Loja)
    Psilodon xerophilicum Martinez & Reyes-Castillo 1985: 109 Brazil (Bahia)
       
LAMPRIMINAE Macleay 1819: 97
LAMPRIMINI MacLeay 1819: 97
  syn. Streptocerini Kikuta 1986: 131  
  Streptocerus Dejean 1833: 174  
    Streptocerus speciosus Fairmaire 1850: 55 Chile, Argentina
      nom. nud. S. speciosus Dejean 1833: 174  
      syn. S. dejeani Solier 1851:44  
      syn. S. eusticticus Philippi 1864:316  
      syn. S. speciosus regius Kriesche 1922:118  
         
LUCANINAE Latreille 1804: 149
BRASILUCANINI Nikolajev 1999: 171
  Brasilucanus Vulcano & Pereira 1961: 473 info  
    Brasilucanus acomus Ratcliffe 1984: 63 info Brazil, Peru
    Brasilucanus alvarengai Vulcano & Pereira 1961: 475 info Brazil, Guyana
       
CASIGNETINI Reid 1999: 176
  syn. Pholidotini Kikuta 1986: 131 (invalid)  
  Casignetus MacLeay 1819: 98 info  
    syn. Pholidotus MacLeay 1819: 97; not Pholidotus Brisson, 1762  
    syn. Chalcimon Dalman 1824: 1  
    Casignetus humboldti (Gyllenhal in Schönherr 1817: 197) (Lamprima) Brazil
      syn. Pholidotus lepidosus MacLeay 1819: 97  
      syn. C. geotrupoides MacLeay 1819: 98  
    Casignetus lindei (Boileau 1899b: 296) (Pholidotus) Brazil
      syn. Pholidotus lombardi Didier & Seguy 1952: 220  
    Casignetus spixi (Perty 1830: 54) (Chalcimon) Brazil
       
CHIASOGNATHINI Burmeister 1847: 334
  Chiasognathus Stephens 1832: 213  
    syn. Tetropthalma Lesson 1833: 5  
    syn. Bomansodus Chalumeau & Brochier 1995: 20  
    syn. Carmenia Molino-Olmedo 2001: 85  
    syn. Ramirezia Molino-Olmedo 2001: 85  
    syn. Carmeniella Molino-Olmedo 2003: 29  
    syn. Ramireziella Molino-Olmedo 2003: 29  
    Chiasognathus beneshi Lacroix 1978: 250 info Chile
    Chiasognathus grantii Stephens 1832: 214 info Chile, Argentina
      syn. Tetropthalma chiloensis Lesson 1833: 6  
      syn. C. affinis R. Philippi in F. Philippi 1859: 658  
      nom. nud. C. brevidens Germain 1911: 47  
      syn. C. pygmaeus Dallas 1933: 74  
      syn. C. grantii holometallicus Rataj 2012: 34 (including unavailable 'varieties' loc. cit.)  
    Chiasognathus impubis Parry 1870: 68 info Chile
    Chiasognathus jousselinii Reiche 1850: 259 info Chile
    Chiasognathus latreillei Solier 1851: 42 info Chile, Argentina
      syn. C. imberbis R. Philippi in F. Philippi 1859: 657  
      syn. C. reichei Thomson 1862: 407  
    Chiasognathus mniszechii Thomson 1862: 406 info Chile, Argentina
      syn. C. schoenemanni Kriesche 1919: 94  
    Chiasognathus sombrus Paulsen & Smith 2010: 56 info Chile
       
  Sphaenognathus Buquet 1838: 104    
    nom. nud. Orthognatus Dejean 1833: 174  
    nom. nud. Orthognathus Dejean 1837: 193  
    syn. Chiasognathinus Didier in Didier & Séguy 1953: 24  
    syn. Sphaenognathinus Chalumeau & Brochier 1995: 20  
    syn. Chiasornithodus Chalumeau & Brochier 1995: 20  
    Sphaenognathus alticollis Möllenkamp 1912: 302 info Peru, Ecuador
      syn. S. matheusi Bartolozzi, Bomans, & Onore 1992:211  
    Sphaenognathus armatus Parry 1872: 73 Colombia
    Sphaenognathus bellicosus Boileau 1904: 277 Colombia
      syn. S. rehni Moxley 1962: 199  
      syn. S. colombiensis Moxley 1962: 202  
    Sphaenognathus bordoni Brochier & Chalumeau 2000: 253 info Colombia
    Sphaenognathus curvipes Benesh 1948: 45 info Bolivia
    Sphaenognathus feisthamelii (Guérin-Méneville 1838: 287) (Chiasognathus) info Colombia, Ecuador, Venezuela
      syn. S. circumflexus Parry 1874: 367  
      syn. S. canaliculatus Parry 1874: 368  
      syn. S. boileaui Lacroix 1969: 587  
      syn. S. villiersi Lacroix 1969: 589  
      syn. S. subtilis Lacroix 1987: 13  
    Sphaenognathus furumii Lacroix 1990: 11 info Bolivia
    Sphaenognathus garleppi Boileau 1899a: 196 Bolivia
      syn. S. manifestus Jakowleff 1901: 176  
      syn. S. inca Kreische 1922: 116  
      syn. S. cartereti Lacroix 1987: 11  
    Sphaenognathus gaujoni (Oberthür 1885: 118) (Chiasognathus) info Ecuador, Peru
      syn. S. leoni Lacroix 1984: 7  
    Sphaenognathus giganteus Boileau 1911: 428 info Bolivia, Ecuador, Peru
    Sphaenognathus higginsi (Parry 1876: 174) (Chiasognathus) Bolivia
    Sphaenognathus kolbei Kreische 1919: 94 info Ecuador
      syn. S. aequatorius Kreische 1940: 40  
      syn. S. sanchezi Bartolozzi, Bomans, & Onore 1992: 223  
    Sphaenognathus lindenii Murray 1857: 221 Ecuador, Venezuela
      syn. S. murrayi Thomson 1862: 409  
    Sphaenognathus mandibularis Boileau 1911: 426 Colombia
    Sphaenognathus metallescens DeLisle 1967: 521 Colombia
    Sphaenognathus metallifer Bomans & Lacroix 1972: 42 Ecuador
    Sphaenognathus monguilloni Lacroix 1972: 47 info Peru
    Sphaenognathus nobilis Parry 1874: 366 info Venezuela
    Sphaenognathus oberon Kreische 1922: 116 Ecuador
    Sphaenognathus peruvianus (Waterhouse 1869: 18) (Chiasognathus) Peru, Ecuador
      syn. S. wallisi Taschenberg 1870: 178  
    Sphaenognathus prionoides Buquet 1838: 104 info Peru, Colombia
      nom. nud. Orthognatus prionoides Dejean 1833: 174  
      syn. S. bispinosus Didier 1928: 11  
      syn. S. spinosus Didier 1928: 11  
      syn. Orthognathus albofuscus Blanchard 1842: plate 7  
      syn. S. pristis DeLisle 1970: 91  
      syn. S. dentifer Lacroix 1972: 33  
      syn. S. bidentatus Ádám 1986: 15  
    Sphaenognathus pseudocurvipes Baba 2009: 39 Argentina
    Sphaenognathus pubescens Waterhouse 1873: 110 Colombia, Venezuela
      syn. S. villosus Ádám 1986: 16  
    Sphaenognathus rotundatus Lacroix 1987: 12 info Venezuela
    Sphaenognathus signatus Parry 1874: 366 Venezuela
      syn. S. praestans Jakowleff 1901: 77  
      syn. S. hemiphanestus DeLisle 1975: 264  
    Sphaenognathus spinifer Boileau 1904: 284 Colombia
      syn. S. lachaumei Lacroix 1984: 5  
    Sphaenognathus taschenbergi Parry 1874: 368 Venezuela
      syn. S. didieri Lacroix 1972: 37  
    Sphaenognathus xerophilus Bartolozzi & Onore 2006 : 364 Peru
       
LUCANINI Latreille 1804: 234
  syn. Dorcini Parry 1864: 86  
  syn. Cladognathini Parry 1870: 75  
  syn. Rhaetulini Miwa 1931: 323  
     
  Apterocyclus Waterhouse 1871:315  
    Apterocyclus honoluluensisWaterhouse 1871: 315 info USA: Hawaii: Kauai
      syn. Apterocyclus deceptor Sharp in Sharp & Scott, 1908: 405.  
      syn. Apterocyclus feminalisSharp in Sharp & Scott, 1908: 405.  
      syn. Apterocyclus varians Sharp in Sharp & Scott, 1908: 404.  
    Apterocyclus kawaii Paulsen & Hawks 2014: 85 info  
    Apterocyclus munroi Sharp in Sharp & Scott 1908: 403 info  
      syn. Apterocyclus adpropinquans Sharp in Sharp & Scott, 1908: 404.  
    Apterocyclus palmatus Van Dyke 1921: 47 info USA: Hawaii: Kauai
    Apterocyclus waterhousei Sharp in Sharp & Scott 1908: 403 info USA: Hawaii: Kauai
     
  Dorcus MacLeay 1819:195  
    Dorcus brevis (Say 1825: 202) (Lucanus) info Eastern North America
      syn. Dorcus caucasicus Ganglbauer 1886: 81  
      syn. Dorcus nanus Casey 1909: 278  
    Dorcus parallelus (Say 1823: 248) (Lucanus) Eastern North America
      syn. Lucanus voeti Schönherr 1817: 326  
      syn. Lucanus oblongus de Charpentier 1825: 214  
      syn. Dorcus costatus LeConte 1866: 35  
      syn. Dorcus carnochani Angell 1916: 70  
       
  Lucanus Scopoli 1763:1  
    syn. Pseudolucanus Hope in Westwood 1845: 30  
    syn. Hexaphyllus Mulsant 1839: 119  
    Lucanus capreolus (Linnaeus 1763: 391) (Scarabaeus) Eastern North America
      syn. L. dama Fabricius 1775: 2  
      syn. L. trigonus Thunberg 1806: 199  
      syn. L. muticus Thunberg 1806: 205  
      nomen nudum Pseudolucanus capreolus ab. muticus Benesh 1938: 273  
    Lucanus elaphus Fabricius 1775: 2 Eastern North America
      nom. nud. L. elaphus ab. carlengi Angell 1916: 70  
    Lucanus mazama (LeConte 1861: 345) (Dorcus) Western North America
      nom. nud. Pseudolucanus mazama ab. bicostatus Angell 1916: 70  
    Lucanus placidus Say 1825: 202 Eastern North America
      syn. L. lentus Laporte 1840: 171  
      nom. nud. L. rupicapra Dejean 1833: 174  
         
PLATYCERINI Mulsant 1842: 593
  syn. Systenocerini Portevin 1931: 3  
  Platycerus Geoffroy 1762: 62 info  
    syn. Systenocerus Wiese 1883: 93  
    Platycerus cribripennis Van Dyke 1928: 107 info USA: California: coastal range
    Platycerus depressus LeConte 1850: 224 info Northeastern USA & Canada
      unavailable name P. piceus Kirby 1837: 141 (misidentification)  
      nom. nud. P. helopioides Dejean 1836: 194  
    Platycerus marginalis Casey 1897: 639 info Western North America
    nom. nud. P. marginalis ab. dietrichae Benesh 1960: 15  
    Platycerus oregonensis Westwood 1845: cvi info Western North America
      syn. P. coerulescens LeConte 1861: 351  
      syn. P. chalybaeus Casey 1897: 639  
    Platycerus virescens (Fabricius 1775: 817) (Lucanus) info East & Central USA, Canada
      syn. Lucanus quercus Weber 1801: 85  
      syn. L. securidens Say 1823: 249  
      syn. P. angustus Casey 1909: 280  
      syn. P. iowanus Casey 1909: 280  
      syn. P. peregrinus Casey 1909: 280  
      nom. nud. P. scaritoides Sturm 1843: 136  
       nom. nud. P. quercicola Knoch 1868: 961  
         
PLATYCEROIDINI Paulsen and Hawks 2008: 2
  Platyceroides Benesh 1946: 175 info  
    Platyceroides aeneus (Van Dyke 1928: 109) (Platycerus) info Western USA
    Platyceroides agassii (LeConte 1861: 345) (Platycerus) Western USA
      orig.comb. Platycerus agassii LeConte 1861: 345  
      misspell. P. agassizii Parry 1870: 114  
      syn. Platycerus parvicollis Casey 1889: 164  
      syn. Platycerus pacificus Casey 1889: 165  
      syn. Platycerus californicus Casey 1885: 331  
      syn. Platycerus pygmaeus Van Dyke 1946: 88  
      unjust. emend. P. agassizi Van Roon 1910: 46  
   
Platyceroides laticollis (Casey 1914: 373) (Platycerus) info
USA: Oregon
    Platyceroides latus (Fall 1901: 291) (Platycerus) info USA: California
    Platyceroides marshalli Paulsen 2015: 1 info USA: Oregon
    Platyceroides opacus (Fall 1906: 393) (Platycerus) info USA: California
    Platyceroides potax Paulsen 2014: 1 info USA: California
    Platyceroides thoracicus (Casey 1895: 154) (Platycerus) USA: California
    Platyceroides viriditinctus (Benesh 1942: 222) (Platycerus) info USA: Oregon
       
  Platyceropsis Benesh 1946: 195 info  
    Platyceropsis keeni (Casey 1895: 153) (Platycerus) info Western North America
      syn. Platycerus pedicellaris Möllenkamp 1912: 302  
         
SCLEROSTOMINI Benesh 1955: 97
  syn. Lissotini Benesh 1955: 73  
  syn. Scortizini Benesh 1955: 103  
  nom.nud. Neoprosopocoilini Maes 1992: 131 (invalid)  
  Aegognathus Leuthner 1883: 445  
    Aegognathus aguirei Arnaud & Bomans 2007: 2 info Peru
    Aegognathus caledoniaensis Grossi et al. 2003: 249 Brazil
    Aegognathus confusus Arnaud & Bomans 2006a: 4 Peru
    Aegognathus iridicolor Lüderwaldt 1935: 518 Brazil
    Aegognathus leuthneri leuthneri Van der Poll 1886: 153 info Colombia, Ecuador, Peru
    Aegognathus leuthneri damasoi Arnaud & Bomans 2006c: 2 Peru
    Aegognathus similis Arnaud & Bomans 2006a: 5 info Peru
    Aegognathus soulai Arnaud & Bomans 2004: 5 info Peru
    Aegognathus spitzi Ohaus 1929: 155 info Brazil
    Aegognathus waterhousei Leuthner 1883: 445 info Peru
       
  Altitatiayus Weinreich 1960: 74 info  
    Altitatiayus campoi Grossi & Koike 2011: 2 Brazil
    Altitatiayus dulceae (Bomans & Arnaud 2002: 6) (Sclerostomus) Brazil
    Altitatiayus godinhorum (Bomans & Arnaud 1996: 3) (Sclerostomus)info Brazil
    Altitatiayus koikei Grossi in Grossi & Almeida 2011: 599 Brazil
    Altitatiayus rotundatus (Boileau 1902: 43) (Sclerostomus) Brazil
      syn. Sclerognathus zikani Ohaus 1929: 158  
    Altitatiayus ruficollis (Lüderwaldt 1931: 425) (Sclerostomus) Brazil
    Altitatiayus trifurcatus (Grossi & Racca-Filho 2004: 1) (Sclerostomus) Brazil
       
  Andinolucanus Arnaud & Bomans 2006a: 2  
    Andinolucanus inesae Arnaud & Bomans 2006a: 3 Peru
     
  Arnaudius Grossi & Bartolozzi 2011: 387  
    Arnaudius bomansi Grossi & Bartolozzi 2011: 388 Peru
    Arnaudius digennaroi (Arnaud et al. 2008: 4) Peru
    Arnaudius koikei (Arnaud et al. 2007: 5) Peru
     
  Apterodorcus Arrow 1943: 139  
    Apterodorcus bacchus (Hope in Westwood 1845: 26) (Lucanus) Chile, Argentina
      nom. nud. Dorcus chilensis Dejean 1833: 174  
      syn. Dorcus bipunctatus F. Philippi 1859: 656  
    Apterodorcus tristis (Deyrolle in Parry 1870: 95) (Sclerostomus) info Chile
         
  Auxicerus Waterhouse 1883: 387  
    syn. Calodaemon Boileau 1897: 248  
    Auxicerus aethiops Jakowlew 1900: 38 info Bolivia
    Auxicerus multicolor (Boileau 1897: 248) (Calodaemon) info Bolivia
    Auxicerus platyceps Waterhouse 1883: 387 info Peru
       
  Caenolethrus Paulsen & Grossi 2009: 26 info  
    Caenolethrus varasi (Nagel 1932: 115) (Sclerostomus) info Chile
       
  Cantharolethrus Thomson 1862: 411  
    Cantharolethrus azambrei Boileau 1897: 275 (Lucanus) info Ecuador, Peru
    Cantharolethrus buckleyi Parry 1872: 77 Ecuador
    Cantharolethrus elongatus Lacroix 1982:23 Peru
    Cantharolethrus homoderoides Kreische 1928: 116 Costa Rica, Nicaragua
    Cantharolethrus luxerii luxerii (Buquet 1843: 51) (Dorcus) info Colombia, Panama
      syn. Pholidotus reichei Hope 1843: 95  
      syn. C. georgius Thomson 1862: 412  
    Cantharolethrus luxerii inflexus Boileau 1899b: 297 Costa Rica
    Cantharolethrus peruvianus Heller 1918: 74 Bolivia, Peru
    Cantharolethrus steinheili Parry 1875: 51 info Peru
       
  Charagmophorus Waterhouse 1895: 495  
    Charagmophorus lineatus eikoae Nagai 2000: 27 Venezuela
    Charagmophorus lineatus lineatus Waterhouse 1895: 495 Brazil
    Charagmophorus umedai Nagai 1996: 6 Venezuela
       
  Chileistomus Weinreich 1960: 74 info  
    Chileistomus cucullatus (Blanchard 1842: plate 12) (Lucanus) info Chile
      nom nud. Scortizus cornutus Thomson 1862:399  
       
  Erichius Maes 1992: 52  
    syn. Epipedus Solier 1851: 49; nec Epipedus Spinola 1837  
    syn. Calvertius Maes 1992: 53  
    syn. Weinreichiellus Maes 1992: 54  
    syn. Neoepipedus Ozdikmen & Turgut 2006: 145  
    Erichius brevicollis (Benesh 1956: 62) (Pycnosiphorus) Chile
      nom. nud. Sclerognathus brevicollis Germain 1911: 66  
    Erichius caelatus (Blanchard 1842: plate 12) (Lucanus) Chile, Argentina
      syn. Lucanus variolosus Hope in Westwood 1845: 25  
      nom. nud. Sclerostomus circumdatus Deyrolle in Parry 1864: 95  
      nom. nud. Sclerognathus cumingi Hope in Harold 1868: 962  
    Erichius calverti (Nagel 1932: 116) (Sclerostomus) Chile
      syn. Pycnosiphorus ariasi Weinreich 1958: 169  
      nom. nud. Sclerognathus calverti Philippi in Germain 1911: 66  
    Erichius costatus (Benesh 1956: 65) (Pycnosiphorus) Chile
      nom.nud. Sclerognathus costatus Germain 1911: 66  
    Erichius darwinii (Hope 1841: 302) (Dorcus) Chile, Argentina
      syn. Dorcus rufipes Solier 1851: 50  
      syn. Dorcus spinolae Solier 1851: 52  
      syn. Dorcus leiocephalus Solier 1851: 53  
      syn. Dorcus modestus F. Philippi 1859: 657  
      unjust. emend. Dorcus liocephalus Van Roon 1910: 45  
    Erichius fairmairii fairmairii (Parry 1864: 61) (Sclerostomus) Chile
    Erichius fairmairii arriguttii (Weinreich 1958: 172) (Pycnosiphorus)  
    Erichius fasciatus (Germain 1855: 397) (Dorcus) Chile
    Erichius femoralis (Guérin-Méneville 1839: 303) (Dorcus) Chile, Argentina
      syn. Dorcus rufifemoralis Curtis 1845: 456  
      syn. Lucanus rubripes Hope in Westwood 1845: 26  
      syn. Dorcus pallidocinctus Fairmaire & Germain 1861: 2  
    Erichius franzae (Weinreich 1958: 173) (Pycnosiphorus) Chile
    Erichius virgatus (Nagel 1928: 290) (Sclerognathus) Chile
    Erichius vittatus vittatus (Eschscholtz 1822: 9) (Lucanus) Chile
      syn. Lucanus rubrovittatus Blanchard 1842: Plate XII, Figure 9  
      syn. Dorcus rouleti Solier 1851: 53  
    Erichius vittatus magnificus Benesh 1956: 60 Chile
    Erichius zabulostictus Paulsen 2010: 104 Chile
       
  Incadorcus Arnaud & Bomans 2006a: 3  
    Incadorcus ashaninka Grossi 2011: 4 Peru
    Incadorcus ayamaranus Grossi 2011: 6 Bolivia
    Incadorcus cuzcoensis Arnaud et al. 2007: 6 Peru
    Incadorcus damasoi Arnaud & Bomans 2006c: 2 Peru
    Incadorcus michelleae Arnaud & Bomans 2006b: 2 Peru
      syn. Incadorcus kirchneri Schenk 2006: 3  
    Incadorcus paulseni Grossi 2011: 9 Bolivia
    Incadorcus shaunai Arnaud et al. 2007: 4 Peru
    Incadorcus zugeri Arnaud & Bomans 2006a: 4 Peru
       
  Leptinopterus Hope 1838: 316  
    syn. Psalicerus Dejean 1833: 174  
    syn. Psalidostomus Burmeister 1847: 377  
    Leptinopterus affinis Parry 1872: 80 Brazil
      nom. nud. L. pulchellus Parry 1864: 85  
      nom. nud. Leptinopterus nitidus ab. lepidus Lüderwaldt 1930: 124  
    Leptinopterus asketus Grossi 2009: 37 Brazil
    Leptinopterus assimilis Grossi 2009: 40 Brazil
    Leptinopterus atramentarius Nagel 1928: 275 Brazil
    Leptinopterus bicolor Lüderwaldt 1931: 423 Brazil
    Leptinopterus burmeisteri Arrow 1943: 135 Brazil
      syn. Psalidostomus polyodontus Burmeister 1847: 381  
      syn. L. nigra Nagel 1924: 401  
      syn. L. unicolor Lüderwaldt 1935: 503  
    Leptinopterus consimilis Möllenkamp 1900: 47 Brazil
    Leptinopterus constricticollis Heller 1924: 63 info Brazil
    Leptinopterus elegans Jakowleff 1900: 632 Brazil
    Leptinopterus erythrocnemus (Burmeister 1847: 378) (Psalidostomus) Argentina, Brazil
      syn. Lucanus tibialis Klug 1825: 431  
      syn. femoratus Thomson 1862: 397  
    Leptinopterus femoratus (Olivier 1789: 17) (Lucanus) info Brazil, Argentina
      nom. nud. Psalicerus erythrocnemus Dejean 1833:174  
      nom. nud. Lucanus rufifemoratus Hope in Westwood 1845:5  
      syn. Leptinopterus lucidus Lüderwaldt 1930: 125  
    Leptinopterus fraternus Westwood 1874: 359 Brazil
    Leptinopterus fryi (Parry 1862: 112) (Psalidostomus) Brazil
    Leptinopterus gracilipes Didier 1928: 80 Brazil
    Leptinopterus gracilis Boileau 1899a: 196 Brazil
      nom. nud. Leptinopterus rufipes Lüderwaldt 1930: 126  
      nom. nud. L. elegans ab. catharinensis Lüderwaldt 1935: 490  
    Leptinopterus ibex (Billberg 1820: 382) (Lucanus) info Brazil
      nom. nud. Psalicerus aries Dejean 1833:174  
      syn. Lucanus sarcorhamphus Laporte 1840: 172  
      syn. Lucanus polyodontus Hope in Westwood 1845: 15  
      nom. nud. Lucanus complanatus Hope in Westwood 1845:5  
      nom. nud. Lucanus wilsoni Hope in Westwood 1845:5  
      syn. Lucanus politus Hope in Westwood 1845: 5  
    Leptinopterus inaharai Lacroix 1982: 26 Brazil
    Leptinopterus mazama Kreische 1926: 384 Brazil
    Leptinopterus melanarius (Hope in Westwood 1845: 15) (Lucanus) Brazil
      syn. Lucanus funereus Hope in Westwood 1845: 15  
      syn. Psalidostomus morio Burmeister 1847: 379  
    Leptinopterus nigrotibialis Lüderwaldt 1935: 506 Brazil
    Leptinopterus nitidus Lüderwaldt 1930: 123 Brazil
    Leptinopterus paranensis Parry 1872: 80 Brazil
    Leptinopterus pellitomarginatus Lüderwaldt 1930: 125 Brazil
    Leptinopterus puncticollis Lüderwaldt 1930: 123 Brazil
    Leptinopterus robustus Lüderwaldt 1930: 127 Brazil
    Leptinopterus suturalis Lüderwaldt 1930: 125 Brazil
    Leptinopterus tibialis (Eschscholtz 1822: 5) (Lucanus) info Argentina, Brazil, Paraguay
      syn. L. femoratus Thomson 1862: 397  
      nom. nud. L. ochraceus Hope MS in Gemminger & Harold 1868: 954  
      nom. nud. L. ochropterus Hope MS in Gemminger & Harold 1868: 954  
      nom. nud. L. ochropterus Hope MS in Boileau 1913: 237  
      nom. nud. L. tibialis ab. fulvus Lüderwaldt 1930: 125  
      nom. nud. L. tibialis ab. castaneus Lüderwaldt 1930: 125  
      nom nud. L. tibialis var. bucki Lüderwaldt 1935: 493  
      syn. L. rotundicollis Lüderwaldt 1930: 124  
      syn. L. luederwaldti de Moraes in Lüderwaldt 1935: 506  
    Leptinopterus vestitus Benesh 1937: 111 Brazil
    Leptinopterus v-nigrum (Hope in Westwood 1845: 15) (Lucanus) info Brazil
      syn. Psalidostomus triangularis Burmeister 1847: 380  
      nom. nud. L. v-nigrum ab. immaculatus Lüderwaldt 1930: 123  
    Leptinopterus ypirangensis Lüderwaldt 1930: 126 Brazil
       
  Macrocrates Burmeister 1847: 381  
    Macrocrates australis Lüderwaldt 1934: 388 Brazil
      syn. M. bucephala Didier 1933: 25  
    Macrocrates bucephalus (Westwood 1845: 15) (Lucanus) Brazil
      nom. nud. Psalicerus nigripes Dejean 1833: 174  
      syn. Lucanus longicornis Burmeister 1848: 382  
      syn. M. rotundicollis Boileau 1913: 237  
      syn. M. formosus Didier 1926: 83  
    Macrocrates galantorum Fujii 1845: 15 Brazil
    Macrocrates invenireus Fujii 1845: 15 Brazil
         
  Metadorcinus Kreische 1922: 126  
    syn. Beneshius Weinreich 1960: 52  
    Metadorcinus amuelleri (Weinreich 1963: 211) (Beneshius) Brazil
    Metadorcinus auritus Kreische 1922: 126 Brazil
    Metadorcinus beneshi (Martínez 1953: 43) (Sclerostomus) Colombia, Ecuador, Peru
    Metadorcinus buckleyi (Waterhouse 1886: 497) (Sclerostomus) Ecuador
    Metadorcinus cruentus (Burmeister 1847: 425) (Sclerostomus) Brazil
      syn. Scortizus cribratus Thomson 1862: 429  
      syn. Sclerostomus tristis Lüderwaldt 1935: 558  
    Metadorcinus dentifer (Möllenkamp 1912: 301) (Scortizus) Brazil
    Metadorcinus ditomoides (Westwood 1855: 208) (Scortizus) Brazil
    Metadorcinus lineatus (Deyrolle 1864: 319) (Sclerostomus) Peru
    Metadorcinus neotragus (Westwood 1855: 207) (Scortizus) Brazil
    Metadorcinus plagiatus (Burmeister 1847: 425) (Sclerostomus) Brazil
      syn. Sclerostomus hastatus Westwood 1855: 205  
    Metadorcinus ranki Grossi & Vaz-de-Mello 2007: 50 Brazil
    Metadorcinus securiformis (Lüderwaldt 1934: 389) (Sclerostomus) Brazil
    Metadorcinus signatipennis (Deyrolle 1864: 319) (Sclerostomus) Brazil
    Metadorcinus sylviae Boucher (1993: 420) (Beneshius) Bolivia
    Metadorcinus tucumanus (Nagel 1932: 119) (Sclerostomus) Argentina
    Metadorcinus yamauchii Arnaud et al. 2008: 4 Peru
       
  Metadorcus Parry 1870: 88  
    syn. Colocephalaria Deyrolle 1864: 317  
    Metadorcus ebeninus Deyrolle 1864: 317 (Platycerus) Brazil or Peru
    Metadorcus rotundatus Parry 1862: 112 (Psalidostomus) Brazil, Uruguay
      nom. nud. M. pachygnathus Parry 1864: 86  
       
  Montesinus Grossi 2016: 219  
    Montesinus machadoi Grossi 2016: 228 Brazil
    Montesinus monnei Grossi 2016: 226 Brazil
    Montesinus tatula Grossi 2016: 227 Brazil
     
  Onorelucanus Bartolozzi & Bomans 1989 : 53  
    syn. Pseudoscortizus Arnaud et al. 2008: 2  
    Onorelucanus aequatorianus Bartolozzi & Bomans 1989: 55 Ecuador
    Onorelucanus boileaui (Weinreich 1960: 81)(Sclerostomus) Peru
    Onorelucanus incredibilis (Arnaud et al. 2008: 3) Peru
    Onorelucanus marujae (Arnaud et al. 2008: 5)(Sclerostomus) Peru
    Onorelucanus nigrosuturalis (Weinreich 1963: 196)(Aegognathus) Ecuador
    Onorelucanus noguchii (Arnaud & Bomans 2007: 3)(Sclerostomus) Peru
    Onorelucanus onorei Lacroix & Bartolozzi 1989: 2 Ecuador
    Onorelucanus pulverosus (Westwood 1875: 243)(Scortizus) Ecuador, Colombia?
      syn. Scortizus ismaeli Fukinuki 2004: 32  
    Onorelucanus rufolineatus (Nagel 1928: 257)(Metadorcus) Ecuador
       
  Pycnosiphorus Solier 1851: 56  
    Pycnosiphorus lessonii lessonii (Buquet 1842: 283) (Lucanus) Chile
    Pycnosiphorus lessonii elongatus (Deyrolle in Parry 1870: 96) (Sclerostomus) Chile
    Pycnosiphorus lessonii marginipennis Deyrolle in Parry 1870: 95 Chile, Argentina
      syn. Pycnosiphorus pulcher Landin 1955: 4  
    Pycnosiphorus lessonii mandibularis (Solier 1851: 56) (Dorcus) Chile
    syn. Pycnosiphorus germaini Numhauser 1985: 71  
    Pycnosiphorus lessonii ugartei Paulsen 2010: 122 Chile
    Pycnosiphorus philippi (Westwood 1864: 61) (Sclerostomus) Chile
      syn. Pycnosiphorus westwoodi Numhauser 1985: 71  
       
  Sclerostomulus Weinreich 1960: 85 info  
    Sclerostomulus nitidus (Benesh 1955: 99) (Sclerostomus) info Chile
    Sclerostomulus tuberculatus (Solier 1851: 54) (Dorcus) info Chile
       
  Sclerostomus Burmeister 1847: 423  
    syn. Sclerognathus Westwood 1845: 26  
    syn. Hemicardanus Heller 1927: 213  
    syn. Sclerostominus Weinreich 1960: 85  
    subgenus Stomosclerus Weinreich 1960: 82  
    Sclerostomus bartolozzii Arnaud & Bomans 2007: 3 Peru
    Sclerostomus costatus (Westwood 1845: 27) (Sclerognathus) Argentina, Brazil
      nom. nud. Psalicerus cuniculus Dejean 1833: 1874  
      syn. Scortizus cuniculus Thomson 1862: 429  
      syn. Hemicardanus interruptocarinulatus Heller 1927: 213  
    Sclerostomus damasoi Arnaud & Bomans 2006c: 3 Peru
    Sclerostomus delislei Weinreich 1961: 433 Brazil
    Sclerostomus gounellei (Boileau 1899b: 299) (Scortizus) info Brazil
      syn. Sclerostomus denticulatus Lüderwaldt 1934: 388  
    Sclerostomus sulcicollis (Möllenkamp 1912: 301) (Scortizus) info Argentina, Brazil
      syn. Sclerostomus costatus genalis Lüderwaldt 1935: 541  
    Sclerostomus truncatus Lüderwaldt 1935: 551 Argentina, Brazil, Paraguay
    Sclerostomus wendyae Arnaud & Bomans 2006b: 3 Peru
      syn. Sclerostomus kirchneri Schenk 2006: 1  
       
  Scortizus Westwood 1834: 119  
    Scortizus maculatus (Klug 1825: 432) (Lucanus) Brazil
      syn. Pholidotus irroratus Hope 1835: 100  
    Scortizus prodigiosus Arnaud et al. 2007: 2 Peru
    Scortizus signatus Weinreich 1963: 206 Bolivia
    Scortizus zischkai Martínez 1957: 225 Bolivia
       
  Zikanius Grossi & Paulsen 2009: 33  info  
    Zikanius aurocinctus (Boileau 1899b: 298)(Sclerostomus) aurocinctus Brazil
      syn. Sclerostomus marginivillosus Lüderwaldt 1935: 535  
       

 

REFERENCES

Benesh, B. 1960. Coleopterorum Catalogus Supplementa, pars 8: Lucanidae. W. Junk, Gravenhage, Netherlands. 178 pp.

Didier, R. and E. Séguy. 1953. Catalogue illustré des lucanides du globe, Texte. Encyclopédie Entomologique (series A) 27: 1-223.

Fujita, H. 2010. The Lucanid beetles of the world. Mushi-Sha's iconographic series of insects, 6. 472 pp. + 248 plates.

Grossi, P.C. 2011. Review of Incadorcus Arnaud & Bomans with the description of three new species from the Yungas of Peru and Bolivia (Coleoptera: Lucanidae: Lucaninae) Zootaxa 2750: 1–14.

Grossi, P.C., and M.J. Paulsen. 2009. Generic limits in South American stag beetles: taxa currently misplaced in Sclerostomus Burmeister (Coleoptera: Lucanidae: Lucaninae: Sclerostomini). Zootaxa 2139: 23-42.

Holloway, B.A. 1960. Taxonomy and phylogeny in the Lucanidae (Insecta: Coleoptera). Records of the Dominion Museum 3: 321-365.

Holloway, B.A. 1968. The relationship of Syndesus MacLeay and Sinodendron Schneider (Coleoptera: Lucanidae). New Zealand Journal of Science 11: 264-269.

Holloway, B.A. 1969. Further studies on generic relationships in Lucanidae (Insecta: Coleoptera) with special reference to the ocular canthus. New Zealand Journal of Science 12: 958-977.

Holloway, B.A. 1997. Elytral surface structures as indicators of relationships in stag beetles, with special reference to the New Zealand species (Coleoptera: Lucanidae). New Zealand Journal of Zoology 24: 47-64.

Holloway, B.A. 1998. A re-evaluation of the genera of New Zealand aesaline stag beetles (Coleoptera: Lucanidae). Journal of the Royal Society of New Zealand 28: 641-656.

Howden, H. and Lawrence, J.H. 1974. The New World Aesalinae, with notes on the North American lucanid subfamilies (Coleoptera, Lucanidae). Canadian Journal of Zoology 52: 1505-1510.

Kikuta, T. 1986. On the higher taxa of the stag beetle family Lucanidae, pp. 131-138. In: J. Aoki (editor). Papers on Entomology Presented to Professor Takeshiko Nakane in Commemoration of His Retirement. Japanese Society of Coleopterology, Tokyo. 277 pp.

Krajcik, M. 2001. Lucanidae of the World, Catalogue – Part 1, Checklist of the stag beetles of the world (Coleoptera: Lucanidae). M. Krajcik, Most, Czech Republic. 108 pp.

Lawrence, J.F. and E.B. Britton. 1991. Coleoptera (Beetles). pp. 543-683. In: I. Naumann, (editor). The Insects of Australia. A Textbook for Students and Research Workers, 2nd Edition Vol. II. (CSIRO) Melbourne University Press, Melbourne. 1137 pp.

Maes, J.-M. 1992. Lista de los Lucanidae (Coleoptera) del mundo. Revista Nicaraguense de Entomologia No. 22: 1-121.

Paulsen, M.J. 2010. The stag beetles of southern South America (Coleoptera: Lucanidae). Bulletin of University of Nebraska State Museum 24: 1-148. Available here.

Smith, A.B.T. 2006. A review of the family-group names for the superfamily Scarabaeoidea (Coleoptera) with corrections to nomenclature and a current classification. Coleopterists Bulletin 60: 144-204.

Triplehorn, C.A. & N.F. Johnson. 2005. Borror and DeLong's Introduction to the Study of Insects, 7th edition. Thomson Brooks/Cole, Belmont, CA. 864 pp.

White, R.E. 1983. A Field Guide to the Beetles of North America. Houghton Mifflin, Boston, MA. 368 pp

....Entomology Home Research UNSM Entomology Database Scarab Workers

Author: M.J. Paulsen - mpaulsen@unlserve.unl.edu
Generated on: 01/JUL/2005
.....Last modified: 17/FEB/2016
UNL State Museum - Division of Entomology