Karla E. Villatoro

villatoro.jpg (32817 bytes)
Photo:  "Scarabs watchout!" Karla Villatoro at Yasuni National  Park, Ecuador, August 1998.
Photo by Andrew Smith.

Karla Villatoro finished her M.S. degree in our lab in 2000. She now lives in Redondo Beach, California.

For her research she revised the genus Trizogeniates (Scarabaeidae: Rutelinae: Geniatini), which was published in Entomotropica in 2002.


Karla Villatoro




Villatoro, K.  2002. Revisión del género neotropical Trizogeniates Ohaus
(Coleoptera: Scarabaeidae: Rutelinae: Geniatini).  Entomotropica 17: 225-294.

Villatoro, K. and M.L. Jameson. 2001. Xenogeniates, a new and unusual genus
of geniatine scarab (Coleoptera: Scarabaeidae: Rutelinae: Geniatini) from Brazil. Annals of the Entomological Society of America 94(6):866-87.

Villatoro, K. 2000. Revision of the genus Trizogeniates (Scarabaeidae: Rutelinae: Geniatini). M. S. Thesis. Department of Entomology, University of Nebraska.

Villatoro, K. 2000. Overview of the tribe Geniatini. URL: http://www-museum.
unl.edu /research/entomology/Guide/Scarabaeoidea/Scarabaeidae/Rutelinae/

Villatoro, K. and M.L. Jameson. 2000. Key to the genera of Geniatini. URL: http://www-Mseum.unl.edu/research/entomology/Guide/Scarabaeoidea
/Scarabaeidae/ Rutelinae/ Rutelinae-Tribes/Geniatini/Geniatini-Key/

Villatoro, K. 2000. Profiles of the species of Trizogeniates. URL: http://www-museum.unl.edu/research/entomology/Guide/Scarabaeoidea/Scarabaeidae/
Rutelinae/ Rutelinae-Tribes/Geniatini/Geniatini-generic-pages/Trizogeniates/

Villatoro, K. 1999-2000. Catalog of the Geniatini. URL: http://www-museum.unl.edu/research/entomology/Guide/Scarabaeoidea/
Scarabaeidae/Rutelinae/ Rutelinae-Tribes/Geniatini/Geniatini-Catalog

Villatoro, K. 1999. Monographic research on Trizogeniates and relationships
of the tribe Geniatini (Rutelinae). URL: http://museum.unl.edu/research/
entomology /obj-triz.html

Villatoro, K. and A. B. T. Smith. 1998. Field Research in Ecuador:
13 July- 7 August 1998. URL: Http://museum.unl.edu/research/

Villatoro, K. 1998. Evaluacion del metodo de PCR para el estudio taxonomico
de Passalidae (Coleoptera: Scarabaeoidea). Licenciatura Thesis. Departamento
de Biologia, Universidad del Valle de Guatemala.

Villatoro, K. 1997. Evaluacion del metodo de PCR para el estudio taxonomico
de Passalidae (Coleoptera:Scarabaeoidea). Paper given at the Latinamerican
reunion of Scarabeoidology, Instituto de Ecologia, Xalapa, Veracruz, Mexico, October 1997.


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University of Nebraska-Lincoln State Museum - Division of Entomology